| Budget Amount *help |
¥18,430,000 (Direct Cost: ¥15,100,000、Indirect Cost: ¥3,330,000)
Fiscal Year 2009: ¥4,810,000 (Direct Cost: ¥3,700,000、Indirect Cost: ¥1,110,000)
Fiscal Year 2008: ¥4,810,000 (Direct Cost: ¥3,700,000、Indirect Cost: ¥1,110,000)
Fiscal Year 2007: ¥4,810,000 (Direct Cost: ¥3,700,000、Indirect Cost: ¥1,110,000)
Fiscal Year 2006: ¥4,000,000 (Direct Cost: ¥4,000,000)
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| Research Abstract |
Photoperiodism is an important biological mechanism and involved in agriculture production but its molecular mechanism remains obscure.
This study analyzed the structures, localization and dynamics of the elements involved in photoreception, circadian oscillation and endocrine switch to reconstruct the photoperiodic system. Comparative studies using different insects revealed a considerable flexibility in the circadian pacemaker structure and photoreceptor regarding to ontogenetic stages and phylogenetic lines. The important factor mediating photpoperiodic system seems to be the indolaminergic pathway, since the upstream regulatory region of N-acetyltransferase(NAT) that catalyze the production of melatonin has several E-boxes where transcription factors CLK/CYK binds, the cells secreting prothoracicotropic horone (PTTH) express melatonin receptor and all PER containing cells have NAT, and melatonin content and NAT transcript level show a circadian oscillation. These cells have the extraretinal photoreception.
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