| Budget Amount *help |
¥310,960,000 (Direct Cost: ¥239,200,000、Indirect Cost: ¥71,760,000)
Fiscal Year 2021: ¥51,090,000 (Direct Cost: ¥39,300,000、Indirect Cost: ¥11,790,000)
Fiscal Year 2020: ¥51,090,000 (Direct Cost: ¥39,300,000、Indirect Cost: ¥11,790,000)
Fiscal Year 2019: ¥51,090,000 (Direct Cost: ¥39,300,000、Indirect Cost: ¥11,790,000)
Fiscal Year 2018: ¥51,090,000 (Direct Cost: ¥39,300,000、Indirect Cost: ¥11,790,000)
Fiscal Year 2017: ¥106,600,000 (Direct Cost: ¥82,000,000、Indirect Cost: ¥24,600,000)
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| Outline of Final Research Achievements |
The present study is performed in the purpose of elucidating the mechanisms of photosynthetic water-oxidation and light-energy absorption, transfer within photosystem (PS)-light harvesting protein (LHC) supercomplexes from various photosynthetic organisms. We analyzed the structures of multiple reaction intermediates of water-oxidation by a pump-probe approach using X-ray free electron laser, and found that an oxygen atom O6 is inserted into the vicinity of O5 within the Mn4CaO5 cluster, the catalyst for water-oxidation, at S3-state generated by 2 flash illumination. This enables formation of an oxygen molecule between O5 and O6. We also analyzed the structures of various PS-LHCs from various organisms, and revealed various light energy transfer pathways in these organisms as well as the changes of PS-LHC supercomplexes occurred during evolution, which provide important insights into the evolution of photosynthetic systems.
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