| Project/Area Number |
01540575
|
|---|
| Research Category |
Grant-in-Aid for General Scientific Research (C)
|
| Allocation Type | Single-year Grants |
|---|
| Research Field |
植物形態・分類学
|
|---|
| Research Institution | University of Tokyo |
|---|
Principal Investigator |
KATO Masahiro Faculty of Science, University of Tokyo Associate Professor, 理学部, 助教授 (20093221)
|
|---|
| Project Period (FY) |
1989 – 1990
|
| Project Status |
Completed (Fiscal Year 1990)
|
| Budget Amount *help |
¥1,900,000 (Direct Cost: ¥1,900,000)
Fiscal Year 1990: ¥400,000 (Direct Cost: ¥400,000)
Fiscal Year 1989: ¥1,500,000 (Direct Cost: ¥1,500,000)
|
|---|
| Keywords | angiosperm / progymnosperm / Ophioglossaceae / carpel / ovule / leaf hyponasty / secondary vascular tissue / pedomorphosis / 下偏生長 / 幼形成熟 / 比較形態学 / 形態形成 / 相同性 / グロッソプテリス |
|---|
| Research Abstract |
1. In order to clarify the anatomical nature of the anomalous vascular tissue of Helminthostachys leylanica, a comparative study was made in branched rhizomes with destroyed shoot Spices. In a portion below the shoot apex to an insertion position of a lateral branch, there are parenchymatous cells and tracheary elements which are radially arranged outside of the primary xylem. These tissue is suggested to be comparable to the secondary vascular tissue of Botrychium and be produced from meristematic cells which appear temporarily in relation to destruction of shoot apices and formation of lateral branches. This evidence supports the idea that the Ophioglossaceae is a living progymnosperm.
2. The origin of angiosperm carpels was examined by comparing them to ophioglossoid fertile leaves. The two organs are similar in the three dimensional architecture, i. e. ovules and sporophores on the adaxial side of carpellary walls and trophores, respectively. Developmental anatomical evidence of ovules accumulated so far suggests that the development of ovules are equivalent to that of leaves. I proposed the Ophioglossoid model in which the angiosperm carpel was derived by pedomorphosis from a three-dimensional leafy organ like ophioglossoid fertile leaves and ovular anatropy is comparable to leaf hyponasty. In this model the fundamental architecture of the carpel had been established prior to carpel enclosure.
|
