| Budget Amount *help |
¥17,550,000 (Direct Cost: ¥13,500,000、Indirect Cost: ¥4,050,000)
Fiscal Year 2018: ¥4,680,000 (Direct Cost: ¥3,600,000、Indirect Cost: ¥1,080,000)
Fiscal Year 2017: ¥5,070,000 (Direct Cost: ¥3,900,000、Indirect Cost: ¥1,170,000)
Fiscal Year 2016: ¥7,800,000 (Direct Cost: ¥6,000,000、Indirect Cost: ¥1,800,000)
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| Outline of Final Research Achievements |
Photosynthetic organisms survive by controlling the photosynthesis and reproduction by detecting deficiency of inorganic nutrients. We found factors involved in this environmental acclimation from the green alga Chlamydomonas. 1) Induction of the CO2-concentrating mechanism (CCM) essential for the maintenance of photosynthesis requires translocation of the post-transcriptional calcium sensor protein CAS into the pyrenoid. 2) A starch sheath around the "pyrenoid" of CO2-fixing enzymes is required for CCM, separating the compartments for CO2-fixing and O2-oxygen evolution. 3) In nitrogen starvation, DYRK-type protein phosphatase TAR1 promotes ROS generation, induces degradation of photosynthesis-related proteins and gametogenesis, and leads to lipid accumulation. 4) Autophagy is required for starch accumulation and maintenance by photosynthesis.
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