| Budget Amount *help |
¥26,260,000 (Direct Cost: ¥20,200,000、Indirect Cost: ¥6,060,000)
Fiscal Year 2019: ¥6,890,000 (Direct Cost: ¥5,300,000、Indirect Cost: ¥1,590,000)
Fiscal Year 2018: ¥8,840,000 (Direct Cost: ¥6,800,000、Indirect Cost: ¥2,040,000)
Fiscal Year 2017: ¥10,530,000 (Direct Cost: ¥8,100,000、Indirect Cost: ¥2,430,000)
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| Outline of Final Research Achievements |
Plants sense the gravity vector and change their growth orientation accordingly, which is known as gravitropism. Based on the Starch-statolith hypothesis, sedimentation of the starch-filled organelle amyloplast in root columella cells or shoot endodermal cells is supposed to trigger gravity sensing in Arabidopsis. The Cholodny-Went model predicts that redistribution (polar transport) of the plant hormone auxin causes differential growth resulting in tropistic responses in roots and shoots. However, little is known about the mechanisms linking the two critical models: Starch-statolith hypothesis and Cholodny-Went model. Using novel imaging techniques (wide-field fluorescent microscope, confocal laser-scanning microscope equipped with optical tweezer and centrifuge fluorescent microscope), we propose a model that gravity-dependent movements of the amyloplasts directly/indirectly facilitate intracellular polar trafficking of the auxin efflux carrier PIN in shoot endodermal cells.
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