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Exploring the socio-cognitive abilities of horses as companion animals: a multi-hierarchical approach from individuals to groups

Research Project

Project/Area Number 18K18342
Research Category

Grant-in-Aid for Early-Career Scientists

Allocation TypeMulti-year Fund
Review Section Basic Section 90030:Cognitive science-related
Research InstitutionTeikyo University of Science & Technology (2021-2022)
Kyoto University (2018-2020)

Principal Investigator

Ringhofer Monamie  帝京科学大学, 生命環境学部, 講師 (20767339)

Project Period (FY) 2018-04-01 – 2023-03-31
Project Status Completed (Fiscal Year 2022)
Budget Amount *help
¥4,030,000 (Direct Cost: ¥3,100,000、Indirect Cost: ¥930,000)
Fiscal Year 2021: ¥910,000 (Direct Cost: ¥700,000、Indirect Cost: ¥210,000)
Fiscal Year 2020: ¥910,000 (Direct Cost: ¥700,000、Indirect Cost: ¥210,000)
Fiscal Year 2019: ¥1,040,000 (Direct Cost: ¥800,000、Indirect Cost: ¥240,000)
Fiscal Year 2018: ¥1,170,000 (Direct Cost: ¥900,000、Indirect Cost: ¥270,000)
Keywordsウマ / 伴侶動物 / 社会性 / 社会的知性 / 個体~集団レベル / 他者への理解 / 他者との行動調整 / 他者理解 / 行動調整 / 個体~群間レベル / 飼育下・野生下環境 / 社会行動 / 行動の同調 / 動きの同調 / 集団行動 / 同調特性 / 家畜化
Outline of Final Research Achievements

In this study, I studied the social interaction between horses and between humans and horses combining experiments on domestic horses and field observations of feral horses through a multi-hierarchical (from individuals to groups) verification. I have investigated horses’ ability of behavioral coordination between individuals and understanding of others from an evolutionary point of view. This study shows that horses have various relationships within and between groups, and build peaceful and complex societies. Furthermore, it was revealed that they are sensitive to the mental and behavioral states of others, including humans, and flexibly adjust their behavior according to those states. It also showed that they have high socio-cognitive abilities toward humans.

Academic Significance and Societal Importance of the Research Achievements

他者理解や同調特性に関する研究は、ヒトと系統進化的に近縁な霊長類を対象とすることが多かったが、近年ではヒトと社会的に近縁なイヌなどの家畜を対象に進んできた。イヌ同様にヒトと社会関係を築くウマを対象とした本研究成果は、ヒトを含む動物の社会性の発達・進化への理解を深めることに貢献する。また350頭以上のウマを見た目で個体識別し、ドローンを用いた新しい行動観察手法を確立するなど、手法面でも重要な意義を持つ。
さらに本研究から得られた知見は、ウマが本来もつ社会性への理解を深めるとともに、それを飼育下ウマの管理に生かすことで、今後のウマのウェルフェア向上およびヒトとウマとのより良い共生の実現に貢献する。

Report

(6 results)
  • 2022 Annual Research Report   Final Research Report ( PDF )
  • 2021 Research-status Report
  • 2020 Research-status Report
  • 2019 Research-status Report
  • 2018 Research-status Report
  • Research Products

    (96 results)

All 2023 2022 2021 2020 2019 2018 Other

All Int'l Joint Research (8 results) Journal Article (27 results) (of which Int'l Joint Research: 11 results,  Peer Reviewed: 14 results,  Open Access: 10 results) Presentation (56 results) (of which Int'l Joint Research: 35 results,  Invited: 1 results) Book (1 results) Remarks (4 results)

  • [Int'l Joint Research] フランス国立農業研究所(INRAE)/ソルボンヌ第3大学(フランス)

    • Related Report
      2021 Research-status Report
  • [Int'l Joint Research] ポルト大学CIBIO/コインブラ大学(ポルトガル)

    • Related Report
      2021 Research-status Report
  • [Int'l Joint Research] ポルト大学CIBIO/コインブラ大学(ポルトガル)

    • Related Report
      2020 Research-status Report
  • [Int'l Joint Research] フランス国立農業研究所(INRAe)/ソルボンヌ第3大学(フランス)

    • Related Report
      2020 Research-status Report
  • [Int'l Joint Research] ポルト大学CIBIO(ポルトガル)

    • Related Report
      2019 Research-status Report
  • [Int'l Joint Research] フランス国立農業研究所/ソルボンヌ第3大学(フランス)

    • Related Report
      2019 Research-status Report
  • [Int'l Joint Research] フランス国立農業研究所/ソルボンヌ大学(フランス)

    • Related Report
      2018 Research-status Report
  • [Int'l Joint Research] ポルト大学CIBIO(ポルトガル)

    • Related Report
      2018 Research-status Report
  • [Journal Article] Horses form cross?modal representations of adults and children.2023

    • Author(s)
      Jardat P., Ringhofer M., Yamamoto S., Gouyet C., Degrande R., Parias C., Reigner F., Calandreau L., Lansade L.
    • Journal Title

      Animal Cognition

      Volume: 26 Issue: 2 Pages: 369-377

    • DOI

      10.1007/s10071-022-01667-9

    • Related Report
      2022 Annual Research Report
    • Peer Reviewed / Int'l Joint Research
  • [Journal Article] Horses cross?modally recognize women and men.2023

    • Author(s)
      Gouyet C., Ringhofer M., Yamamoto S., Jardat P., Parias C., Reigner F., Calandreau L. & Lansade L.
    • Journal Title

      Scientific Reports

      Volume: 13 Issue: 1 Pages: 3864-3864

    • DOI

      10.1038/s41598-023-30830-6

    • Related Report
      2022 Annual Research Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] Population characteristics of feral horses impacted by anthropogenic factors and their management implications2022

    • Author(s)
      Mendonca R.S., Pinto P., Maeda T., Inoue S., Ringhofer M., Yamamoto S., Hirata S.
    • Journal Title

      Frontiers in Ecology and Evolution

      Volume: 10 Pages: 848741-848741

    • DOI

      10.3389/fevo.2022.848741

    • Related Report
      2022 Annual Research Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] 動物心理・行動・生態学から探るウマの社会性2022

    • Author(s)
      リングホーファー 萌奈美
    • Journal Title

      Hoppophile

      Volume: 91 Pages: 1-12

    • Related Report
      2022 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Horses with sustained attention follow the pointing of a human who knows where food is hidden2021

    • Author(s)
      Monamie Ringhofer, Milena Troesch, Lea Lansade, Shinya Yamamoto
    • Journal Title

      Scientific Reports

      Volume: 11 Issue: 1 Pages: 1-9

    • DOI

      10.1038/s41598-021-95727-8

    • NAID

      120007141733

    • Related Report
      2021 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] Social determinants of affiliation and cohesion in a population of feral horses2021

    • Author(s)
      Renata S. Mendonca, Pinto Pandora, Sota Inoue, Monamie Ringhofer, Raquel Godinho, Staoshi Hirata
    • Journal Title

      Applied Animal Behaviour Science

      Volume: 245 Pages: 105496-105496

    • DOI

      10.1016/j.applanim.2021.105496

    • Related Report
      2021 Research-status Report
    • Peer Reviewed
  • [Journal Article] Aerial drone observations identified a multilevel society in feral horses2021

    • Author(s)
      Maeda Tamao、Ochi Sakiho、Ringhofer Monamie、Sosa Sebastian、Sueur C?dric、Hirata Satoshi、Yamamoto Shinya
    • Journal Title

      Scientific Reports

      Volume: 11 Issue: 1 Pages: 1-12

    • DOI

      10.1038/s41598-020-79790-1

    • NAID

      120006951653

    • Related Report
      2020 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] A mathematical model of herding in horse-harem group2020

    • Author(s)
      Go Clark Kendrick、Ringhofer Monamie、Lao Bryan、Kubo Takatomi、Yamamoto Shinya、Ikeda Kazushi
    • Journal Title

      Journal of Ethology

      Volume: 38 Issue: 3 Pages: 343-353

    • DOI

      10.1007/s10164-020-00656-0

    • Related Report
      2020 Research-status Report
    • Peer Reviewed / Int'l Joint Research
  • [Journal Article] ウマ学ことはじめ 連載第16回「ヒトの心を読むウマ」2020

    • Author(s)
      山本真也、リングホーファー萌奈美
    • Journal Title

      モンキー

      Volume: 4(4) Pages: 98-99

    • Related Report
      2020 Research-status Report 2019 Research-status Report
  • [Journal Article] ウマ学ことはじめ 連載第19回「群れを保つためのメンバーの役割分担」2020

    • Author(s)
      リングホーファー萌奈美
    • Journal Title

      モンキー

      Volume: 5(3) Pages: 70-71

    • Related Report
      2020 Research-status Report
  • [Journal Article] Feral horses’ (Equus ferus caballus) behavior toward dying and dead conspecifics2020

    • Author(s)
      Renata Mendonca, Monamie Ringhofer, Pandora Pinto, Sota Inoue, Satoshi Hirata
    • Journal Title

      Primates

      Volume: 61(1) Issue: 1 Pages: 49-54

    • DOI

      10.1007/s10329-019-00728-x

    • Related Report
      2019 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] Herding mechanisms to maintain the cohesion of a harem group: two interaction phases during herding2020

    • Author(s)
      Monamie Ringhofer, Clark K Go, Sota Inoue, Renata S Mendonca, Sotoshi Hirata, Takatomi Kubo, Kazushi Ikeda, Shinya Yamamoto
    • Journal Title

      Journal of Ethology

      Volume: 38(1) Issue: 1 Pages: 71-77

    • DOI

      10.1007/s10164-019-00622-5

    • Related Report
      2019 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] Estudos preliminares sobre o comportamento e ecologia dos cavalos ferais na Serra d’Arga, Norte de Portugal2020

    • Author(s)
      Renata S Mendonca, Monamie Ringhofer, Sota Inoue, Satoshi Hirata
    • Journal Title

      Atas dos seminários - Percursos do Homem e do Garrano (in Portuguese)

      Volume: -

    • Related Report
      2019 Research-status Report
  • [Journal Article] Horses prefer to solicit a person who previously observed a food-hiding process to access this food: A possible indication of attentional state attribution2019

    • Author(s)
      Milena Trosch, Monamie Ringhofer, Shinya Yamamoto, Julie Lemarchand, Celine Parias, Flore Lormant, Lea Lansade
    • Journal Title

      Behavioural Processes

      Volume: 166 Pages: 103906-103906

    • DOI

      10.1016/j.beproc.2019.103906

    • Related Report
      2019 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] Lateral position preference in grazing feral horses2019

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata S Mendonca, Satoshi Hirata
    • Journal Title

      Ethology

      Volume: 126 Issue: 1 Pages: 111-119

    • DOI

      10.1111/eth.12966

    • Related Report
      2019 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] 野生ウマの生と死: オオカミによる捕食と死の認識2019

    • Author(s)
      リングホーファー萌奈美、 レナータ・メンドンサ
    • Journal Title

      生物の科学 遺伝

      Volume: 5月号 Pages: 237-243

    • Related Report
      2019 Research-status Report
  • [Journal Article] 野生ウマの社会: ポルトガル・アルガ山のガラノ種の事例を中心に2019

    • Author(s)
      平田聡、リングホーファー萌奈美、井上漱太、前田玉青、越智咲穂、山本真也
    • Journal Title

      生物の科学 遺伝

      Volume: 5月号 Pages: 230-237

    • Related Report
      2019 Research-status Report
  • [Journal Article] ウマ学ことはじめ 連載第14回「モンゴルのウマと遊牧」2019

    • Author(s)
      松沢哲郎、リングホーファー萌奈美
    • Journal Title

      モンキー

      Volume: 4(2) Pages: 42-43

    • Related Report
      2019 Research-status Report
  • [Journal Article] Life and death of feral horses: Predation by wolves and horses recognition of death2019

    • Author(s)
      Monamie Ringhofer, Renata S Mendonca
    • Journal Title

      Revista General de erecho Animal y Estudios Interdisciplinares de Bienestrar Animal

      Volume: 4 Pages: 119-131

    • Related Report
      2019 Research-status Report
  • [Journal Article] ウマ学ことはじめ 連載第15回「気ままに生きるスペインのポニー」2019

    • Author(s)
      リングホーファー萌奈美、レナータ・メンドンサ
    • Journal Title

      モンキー

      Volume: 4(3) Pages: 70-71

    • Related Report
      2019 Research-status Report
  • [Journal Article] ウマ学ことはじめ 連載第12回 「野生ウマの死とオオカミ」2019

    • Author(s)
      レナータ・メンドンサ、リングホーファー 萌奈美
    • Journal Title

      モンキー

      Volume: 3(4) Pages: 98-99

    • Related Report
      2018 Research-status Report
  • [Journal Article] 野生ウマの生と死: オオカミによる捕食と死の認識2019

    • Author(s)
      リングホーファー 萌奈美、 レナータ・メンドンサ
    • Journal Title

      生物の科学 遺伝

      Volume: 3 Pages: 237-243

    • Related Report
      2018 Research-status Report
  • [Journal Article] 野生ウマの社会: ポルトガル・アルガ山のガラノ種の事例を中心に2019

    • Author(s)
      平田 聡、リングホーファー 萌奈美、井上 漱太、前田 玉青、越智 咲穂、山本 真也
    • Journal Title

      生物の科学 遺伝

      Volume: 3 Pages: 230-237

    • Related Report
      2018 Research-status Report
  • [Journal Article] Spatial positioning of individuals in a group of feral horses: a case study using drone technology2019

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata S. Mendonca, Carlos Pereira, Satoshi Hirata
    • Journal Title

      Mammal Research

      Volume: 64 Issue: 2 Pages: 249-259

    • DOI

      10.1007/s13364-018-0400-2

    • Related Report
      2018 Research-status Report
    • Peer Reviewed / Open Access / Int'l Joint Research
  • [Journal Article] Feral horses’ (Equus ferus caballus) behavior toward dying and dead conspecifics2019

    • Author(s)
      Renata S Mendonca, Monamie Ringhofer, Pandora Pinto, Sota Inoue, Satoshi Hirata
    • Journal Title

      Primates

      Volume: 印刷中

    • Related Report
      2018 Research-status Report
    • Peer Reviewed / Open Access
  • [Journal Article] 馬と人間2018

    • Author(s)
      リングホーファー 萌奈美
    • Journal Title

      科学

      Volume: 11 Pages: 1125-1126

    • Related Report
      2018 Research-status Report
  • [Journal Article] ウマ学ことはじめ 連載第11回 「オスがあやつる群れの性質」2018

    • Author(s)
      リングホーファー 萌奈美
    • Journal Title

      モンキー

      Volume: 3(3) Pages: 70-71

    • Related Report
      2018 Research-status Report
  • [Presentation] he Role of Intentional Communication in a Cooperative Activity in Horses (Equus caballus)2022

    • Author(s)
      Gema Palacino-Gonzalez, Tamao Maeda, Monamie Ringhofer, Shinya Yamamoto
    • Organizer
      The 17th International Symposium of Primatology and Wildlife Science
    • Related Report
      2022 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Comparing the Social Relationships of Feral Horses during Breeding and Non-breeding Season Using Drones and GPS Tags.2022

    • Author(s)
      Tamao Maeda, Renata Mendonca, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto.
    • Organizer
      The 17th International Symposium of Primatology and Wildlife Science
    • Related Report
      2022 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Horses with sustained attention follow the pointing of a human who knows where food is hidden2022

    • Author(s)
      Monamie Ringhofer, Milena Troesch, Lea Lansade, Shinya Yamamoto
    • Organizer
      日本動物心理学会第82回大会
    • Related Report
      2022 Annual Research Report
  • [Presentation] Effect of Oxytocin on the Inter-species Relationship Between Horse and Human2022

    • Author(s)
      China Kobayashi, Monamie Ringhofer, Kodzue Kinoshita, Kenji Miyata, Shinya Yamamoto
    • Organizer
      日本動物心理学会第82回大会
    • Related Report
      2022 Annual Research Report
  • [Presentation] ウマは自分と似た見た目の個体と仲良くなりやすいか2022

    • Author(s)
      橋本 和佳奈、山本 真理子、リングホーファー 萌奈美
    • Organizer
      日本ウマ科学会第35回大会
    • Related Report
      2022 Annual Research Report
  • [Presentation] オキシトシンが馬と人の種間関係に与える影響2022

    • Author(s)
      小林 知奈、リングホーファー 萌奈美、木下 こづえ、池田 泰彦、宮田 健二、山本 真也
    • Organizer
      日本人間行動進化学会
    • Related Report
      2022 Annual Research Report
  • [Presentation] Comparing the Social Relationships of Feral Horses during Breeding and Non-breeding Season Using Drones and GPS Tags2022

    • Author(s)
      Tamao Maeda, Renata Mendoca, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      The 17th International Symposium of Primatology and Wildlife Science
    • Related Report
      2021 Research-status Report
    • Int'l Joint Research
  • [Presentation] The Role of Intentional Communication in a Cooperative Activity in Horses (Equus caballus)2022

    • Author(s)
      Gema Palacino-Gonzalez, Tamao Maeda, Monamie Ringhofer, Shinya Yamamoto
    • Organizer
      The 17th International Symposium of Primatology and Wildlife Science
    • Related Report
      2021 Research-status Report
    • Int'l Joint Research
  • [Presentation] Mathematical Model of a Horse and the Rider during a Jump2021

    • Author(s)
      Asahi Tsuruo, Monamie Ringhofer, Shinya Yamamoto, Kazushi IkedA
    • Organizer
      APSIPA ASC 2021
    • Related Report
      2021 Research-status Report
    • Int'l Joint Research
  • [Presentation] 「ウマの社会性と社会的認知能力」2021

    • Author(s)
      リングホーファー萌奈美
    • Organizer
      行動遺伝学研究会「家畜化機構の解明」
    • Related Report
      2021 Research-status Report
    • Invited
  • [Presentation] Human-Horse Interaction Model in Jumping2021

    • Author(s)
      鶴尾 明大、リングホーファー 萌奈美、山本 真也、池田 和司
    • Organizer
      HAIシンポジウム2021
    • Related Report
      2020 Research-status Report
  • [Presentation] Examining the possible mechanisms and adaptive significance of single and multi-male groups in horse societies2021

    • Author(s)
      Pandora Pinto, Renata Mendonca, Monamie Ringhofer, Kodzue Kinoshita, Satoshi Hirata
    • Organizer
      The 15th international symposium of primatology and wildlife science
    • Related Report
      2020 Research-status Report
    • Int'l Joint Research
  • [Presentation] ウマはヒトからの情報の信憑性を見分ける:認知能力と注意力との関連性2020

    • Author(s)
      リングホーファー萌奈美、山本真也
    • Organizer
      日本動物行動学会第39回大会
    • Related Report
      2020 Research-status Report
  • [Presentation] Mathematical Model of Horse Riding during Jumping2020

    • Author(s)
      鶴尾明大、リングホーファー萌奈美、山本真也、池田和司
    • Organizer
      計測自動制御学会システム・情報部門学術講演会
    • Related Report
      2020 Research-status Report
  • [Presentation] ウマはヒトから与えられた情報の信憑性を見分けられるか:その実験成績と注意力との関連性2020

    • Author(s)
      リングホーファー萌奈美、山本真也
    • Organizer
      第33回ウマ科学会
    • Related Report
      2020 Research-status Report
  • [Presentation] ドローンから見る野生ウマの重層社会2020

    • Author(s)
      前田玉青、越智咲穂、リングホーファー萌奈美、セバスチャン・ソーサ、セドリック・スール、平田聡、山本真也
    • Organizer
      第33回ウマ科学会
    • Related Report
      2020 Research-status Report
  • [Presentation] Do macho horse have all the mojo? Examining testosterone concentrations in feral stallions2020

    • Author(s)
      Pandora Pinto, Renata Mendonca, Monamie Ringhofer, Kodzue Kinoshita, Satoshi Hirata
    • Organizer
      The 14th International Symposium of Primatology and Wildlife Science
    • Related Report
      2020 Research-status Report
    • Int'l Joint Research
  • [Presentation] Mathematical model of horse and rider interaction during horse jumping2020

    • Author(s)
      Asahi Tsuruo, Monamie Ringhofer, Shinya Yamamoto, Kazushi Ikeda
    • Organizer
      Asia-Pacific Signal and Information Processing Association Annual Summit and Conference 2020
    • Related Report
      2020 Research-status Report
    • Int'l Joint Research
  • [Presentation] Behavioral responses of feral horses toward dying and dead conspecifics2019

    • Author(s)
      Monamie Ringhofer, Renata S Mendonca
    • Organizer
      The 14th International Conference on Environmental Enrichment
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Estimation of force working between individuals in feral horses2019

    • Author(s)
      Sota Inoue, Keiichiro Fukazawa, Shinya Yamamoto, Monamie Ringhofer, Renata S Mendonça, Satoshi Hirata
    • Organizer
      ASAB summer conference 2019
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Multi-level structure in feral horse society: Evidence from the aerial observation from drones2019

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      ASAB Summer Conference 2019
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Collective departure in feral horses2019

    • Author(s)
      Sota Inoue, Colin Torney, Shinya Yamamoto, Monamie Ringhofer, Renata S Mendonca, Satoshi Hirata
    • Organizer
      The 12th international symposium of primatology and wildlife science
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Mechanism to maintain group cohesion: communication of feral horses (Equus Cavallus) living in Serra d’Arga, Portugal2019

    • Author(s)
      Monamie Ringhofer, Renata S Mendonca, Shinya Yamamoto, Satoshi Hirata, Tetsuro Matsuzawa
    • Organizer
      Protolang 6
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Collective departure in feral horses: combining wild animals with drone2019

    • Author(s)
      Sota Inoue S, Colin Torney, Shinya Yamamoto, Monamie Ringhofer, Renata S Mendonca, Satoshi Hirata
    • Organizer
      SWARM2019: The 3rd International Symposium on Swarm Behavior and Bio-inspired Robotics
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Differences in spatial positioning of individuals among feral horse groups2019

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata S Mendonca, Staoshi Hirata
    • Organizer
      JSPS- The 8th international workshop on tropical biodiversity and conservation
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Modelling the Herding of Garrano Horses in the Wild2019

    • Author(s)
      Clark Kendrick Go, Monamie Ringhofer, Bryan Lao, Takatomi Kubo, Shinya Yamamoto, Kazushi Ikeda
    • Organizer
      Society of Mathematical Biology
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Herding mechanisms to maintain the cohesion of a harem group: two interaction phases during herding2019

    • Author(s)
      Monamie Ringhofer, Clark Kendrick C. Go, Sota Inoue, Renata S Mendonca, Satoshi Hirata, Takatomi Kubo, Kazushi Ikeda, Shinya Yamamoto
    • Organizer
      The 2nd International Wild Equid Conference
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Feral horses’ (Equus ferus caballus) behavior toward dying and dead conspecifics2019

    • Author(s)
      Renata S Mendonca, Monamie Ringhofer, Shinya Yamamoto, Satoshi Hirata, Tetsuro Matsuzawa
    • Organizer
      The 2nd International Wild Equid Conference
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Differences in spatial positioning of individuals among feral horse groups2019

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata S Mendonca, Staoshi Hirata
    • Organizer
      The 2nd International Wild Equid Conference
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Multi-level structure in feral horse society: Evidence from the aerial observation from drones2019

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      The 2nd International Wild Equid Conference
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Multi-level structure in feral horse society: Evidence from the aerial observation from drones2019

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      The 12th International Symposium of Primatology and Wildlife Science
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Multi-level structure in feral horse society: Evidence from the aerial observation from drones2019

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      SWARM2019: The 3rd International Symposium on Swarm Behavior and Bio-inspired Robotics
    • Related Report
      2019 Research-status Report
    • Int'l Joint Research
  • [Presentation] Horses take into account whether a human has previously observed a food-hiding event to decide whom to solicit to access this food2019

    • Author(s)
      Milena Trosch, Monamie Ringhofer, Shinya Yamamoto, Julie Lemarchand, Celine Parias, Flore Lormant, Lea Lansade
    • Organizer
      日本動物心理学会第79回大会
    • Related Report
      2019 Research-status Report
  • [Presentation] 野生ウマにおける集団の意思決定過程2019

    • Author(s)
      井上漱太, Colin Torney, 山本真也, リングホーファー萌奈美, Renata S Mendonca, 平田聡
    • Organizer
      第38回日本動物行動学会大会
    • Related Report
      2019 Research-status Report
  • [Presentation] Inter-group relationships of feral horses (Equus caballus) in Serra D’Arga, Portugal2019

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      プリマーテス研究会
    • Related Report
      2018 Research-status Report
  • [Presentation] The objective investigation of interspecies behavioral synchronization in horse riding2019

    • Author(s)
      Sakiho Ochi, Monamie Ringhofer, Shinya Yamamoto
    • Organizer
      プリマーテス研究会
    • Related Report
      2018 Research-status Report
  • [Presentation] Mechanisms of herding to maintaining cohesiveness of harem group: different reaction of females toward the male and other females2019

    • Author(s)
      Monamie Ringhofer
    • Organizer
      The 11th International Symposium on Primatology and Wildlife Science
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Multi-level structure in feral horse society: Evidence from the aerial observation from drones2019

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      The 11th International Symposium on Primatology and Wildlife Science
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Modelling the Herding of Garrano Horses in the Wild2019

    • Author(s)
      Clark Kendrick G, Monamie Ringhofer, Bryan Lao, Takatomi Kubo, Shinya Yamamoto, Kazushi Ikeda
    • Organizer
      Society of Mathematical Biology
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Individual differences in spatial positioning in feral horses2018

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata Mendonca, Satoshi Hirata
    • Organizer
      Conference on Collective behavior
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Differences in spatial positioning of individuals among feral horse groups2018

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata Mendonca, Satoshi Hirata
    • Organizer
      The 55th Annual Conference of the Animal Behavior Society
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Decision making of mares reacting to the stallion’s herding in a harem group of feral horses (Equus caballus)2018

    • Author(s)
      Monamie Ringhofer, Clark Kendrick C Go, Sota Inoue, Renata S Mendonca, Satoshi Hirata, Takatomi Kubo, Kazushi Ikeda, Shinya Yamamoto
    • Organizer
      International Society of Behavioral Ecology 2018 conference
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] How do the departures begin in feral horses groups?2018

    • Author(s)
      Sota Inoue, Colin Torney, Shinya Yamamoto, Monamie Ringhofer, Renata Mendonca, Satoshi Hirata
    • Organizer
      Symposium on Systems Science of Bio-Navigation 2018
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Do feral horses form multilevel society?2018

    • Author(s)
      Tamao Maeda, Sakiho Ochi, Monamie Ringhofer, Satoshi Hirata, Shinya Yamamoto
    • Organizer
      The 10th International Symposium on Primatalogy and Wildlife Science
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Do female horses (Equus caballus) choose their group?2018

    • Author(s)
      Sakiho Ochi, Monamie Ringhofer, Renata S. Mendonca, Shinya Yamamoto, Satoshi Hirata
    • Organizer
      The 10th International Symposium on Primatalogy and Wildlife Science
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Differences in spatial positioning of individuals among feral horse groups2018

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata Mendonca, Satoshi Hirata
    • Organizer
      The 7th International Workshop on Tropical Biodiversity Conservation focusing on Large Animal Studies
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Lateral position preferences in feral horses2018

    • Author(s)
      Sota Inoue, Shinya Yamamoto, Monamie Ringhofer, Renata Mendonca, Satoshi Hirata
    • Organizer
      The 10th International Symposium on Primatology and Wildlife Science
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Do female horses (Equus caballus) choose their group?2018

    • Author(s)
      Sakiho Ochi, Monamie Ringhofer, Renata S Mendonça, Shinya Yamamoto, Satoshi Hirata
    • Organizer
      The 10th International Symposium on Primatalogy and Wildlife Science
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] Horses’ society and males’ behaviour2018

    • Author(s)
      Monamie Ringhofer
    • Organizer
      Percursos do Homem e do Garrano
    • Related Report
      2018 Research-status Report
    • Int'l Joint Research
  • [Presentation] 野生下ウマにおける負傷した仔ウマに対する母親と同群・他群他個体の反応2018

    • Author(s)
      リングホーファー萌奈美、井上漱太、Pandora Pinto、越智咲穂、前田玉青、Renata S. Mendonca
    • Organizer
      日本動物行動学会第37回大会
    • Related Report
      2018 Research-status Report
  • [Presentation] ハレム群におけるハーディング時のメスの反応:空撮と動作追跡を用いた動きの解析2018

    • Author(s)
      リングホーファー萌奈美、Clark K.C.Go、井上漱太、 Renata S.Mendonca、 平田聡、久保孝富、 池田和司、 山本真也
    • Organizer
      第31回ウマ科学会
    • Related Report
      2018 Research-status Report
  • [Presentation] Behavioral responses of a mother and other in-group and out-group individuals toward a wounded foal: a case report in feral horses (Equus Caballus)2018

    • Author(s)
      Monamie Ringhofer, Sota Inoue, Pandora Pinto, Sakiho Ochi, Tamao Maeda, Renata S. Mendonca
    • Organizer
      第78回日本動物心理学会
    • Related Report
      2018 Research-status Report
  • [Presentation] What is the unity of rider and horse: objective evaluation and investigation of effects on their psychological states2018

    • Author(s)
      Sakiho Ochi, Monamie Ringhofer, Shinya Yamamoto
    • Organizer
      第78回日本動物心理学会
    • Related Report
      2018 Research-status Report
  • [Presentation] ウマの群れにおける個体の空間配置の多様性とその要因2018

    • Author(s)
      井上漱太、山本真也、リングホーファー萌奈美、Renata S. Mendonca、平田聡、
    • Organizer
      日本動物行動学会第37回大会
    • Related Report
      2018 Research-status Report
  • [Presentation] 野生ウマ(Equus caballus)の成体メスはなぜ移籍をするのか?-移籍先選択条件からの考察 -2018

    • Author(s)
      越智咲穂、リングホーファー萌奈美、Renata S. Mendonca、山本真也、平田聡
    • Organizer
      日本動物行動学会第37回大会
    • Related Report
      2018 Research-status Report
  • [Presentation] ウマは重層社会をつくるか? ドローンから見る野生ウマの群れ間関係2018

    • Author(s)
      前田玉青、越智咲穂、リングホーファー萌奈美、平田聡、山本真也
    • Organizer
      日本動物行動学会第37回大会
    • Related Report
      2018 Research-status Report
  • [Book] 『はじめての心理学概論』中 「第12章 進化・動物心理学」2019

    • Author(s)
      リングホーファー 萌奈美(著)  古見 文一、小山内 秀和、樋口 洋子、津田 裕之(編)
    • Total Pages
      13
    • Publisher
      ナカニシヤ出版
    • ISBN
      9784779513619
    • Related Report
      2018 Research-status Report
  • [Remarks] 研究に関するホームページ

    • URL

      https://monamieringhofer.wixsite.com/monamieringhofer

    • Related Report
      2022 Annual Research Report
  • [Remarks] 研究紹介ページ

    • URL

      https://monamieringhofer.wixsite.com/monamieringhofer

    • Related Report
      2021 Research-status Report
  • [Remarks] Monamie Ringhofer's web site

    • URL

      https://monamieringhofer.wixsite.com/monamieringhofer

    • Related Report
      2020 Research-status Report 2018 Research-status Report
  • [Remarks] Monamir Ringhofer's web site

    • URL

      https://monamieringhofer.wixsite.com/monamieringhofer

    • Related Report
      2019 Research-status Report

URL: 

Published: 2018-04-23   Modified: 2024-01-30  

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