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Study on site-fidelity and homing behaviour in coastal fish using wideband-frequency ultrasonic telemetry: from individual to population

Research Project

Project/Area Number 25712022
Research Category

Grant-in-Aid for Young Scientists (A)

Allocation TypePartial Multi-year Fund
Research Field Aquatic bioproduction science
Research InstitutionKyoto University

Principal Investigator

MITAMURA Hiromichi  京都大学, 情報学研究科, 准教授 (20534423)

Project Period (FY) 2013-04-01 – 2016-03-31
Project Status Completed (Fiscal Year 2015)
Budget Amount *help
¥24,830,000 (Direct Cost: ¥19,100,000、Indirect Cost: ¥5,730,000)
Fiscal Year 2015: ¥8,060,000 (Direct Cost: ¥6,200,000、Indirect Cost: ¥1,860,000)
Fiscal Year 2014: ¥8,320,000 (Direct Cost: ¥6,400,000、Indirect Cost: ¥1,920,000)
Fiscal Year 2013: ¥8,450,000 (Direct Cost: ¥6,500,000、Indirect Cost: ¥1,950,000)
Keywordsバイオロギング / バイオテレメトリー / 高精度測位 / 群れ / 行動学 / 個体群
Outline of Final Research Achievements

Some aquatic animals have site-fidelity and homing ability to their habitats and spawning sites. In this study, we have monitored site-fidelity and homing behaviours in coastal fishes at the levels of individuals and a fish school. The wideband-frequency ultrasonic telemetry system has been developed for monitoring schooling behaviour. The telemetry system can locate a lot of individuals at the same time. Our results demonstrated forms and volume of a school, interaction behaviour, and school homing/site-fidelity to their habitat in several coastal fish species.

Report

(4 results)
  • 2015 Annual Research Report   Final Research Report ( PDF )
  • 2014 Annual Research Report
  • 2013 Annual Research Report
  • Research Products

    (44 results)

All 2016 2015 2014 2013

All Journal Article (7 results) (of which Peer Reviewed: 7 results,  Open Access: 1 results,  Acknowledgement Compliant: 1 results) Presentation (37 results) (of which Int'l Joint Research: 15 results,  Invited: 1 results)

  • [Journal Article] 溶存酸素量の低下に対する ヒラメ Paralichthys olivaceusの行動変化2016

    • Author(s)
      河野友香、西澤秀明、荒井修亮、三田村啓理.
    • Journal Title

      海洋理工学会誌

      Volume: 印刷中

    • NAID

      130005162860

    • Related Report
      2015 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Pitching stability of diving seabirds during underwater locomotion: a comparison among alcids and a penguin.2016

    • Author(s)
      Takuji Noda, Dale M Kikuchi, Akinori Takahashi, Hiromichi Mitamura, Nobuaki Arai.
    • Journal Title

      Animal Biotelemetry

      Volume: 印刷中

    • NAID

      120005997664

    • Related Report
      2015 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Measurement of activity for sit-and-wait predator, red-spotted grouper, using acoustic acceleration transmitter2015

    • Author(s)
      Nao Yoshida, Hiromichi Mitamura, Hideaki Okamoto, Nobuaki Arai
    • Journal Title

      Journal of Advanced Marine Science and Technology Society

      Volume: 1 Pages: 1-5

    • NAID

      130005087790

    • Related Report
      2015 Annual Research Report
    • Peer Reviewed / Open Access / Acknowledgement Compliant
  • [Journal Article] Measurement of activity for sit-and-wait predator, red-spotted grouper, using acoustic acceleration transmitter2015

    • Author(s)
      Yoshida N, Mitamura H, Okamoto H, Arai N
    • Journal Title

      Journal of Advanced Marine Science and Technology Society

      Volume: 印刷中

    • NAID

      130005087790

    • Related Report
      2014 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Use of a gyroscope/accelerometer data logger to identify alternative feeding behaviours in fish2014

    • Author(s)
      Kawabata Y, Noda T, Nakashima Y, Nanami A, Sato T, Takebe T, Mitamura H, Arai N, Yamaguchi T, Soyano K.
    • Journal Title

      The Journal of Experimental Biology

      Volume: 217 Pages: 3204-3208

    • NAID

      120006987132

    • Related Report
      2014 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Harvesting energy from the oscillation of aquatic animals: testing a vibration-powered generator for bio-logging data logger systems2014

    • Author(s)
      Noda T, Okuyama J, Kawabata Y, Mitamura H, Arai N.
    • Journal Title

      Journal of Advanced Marine Science and Technology Society

      Volume: 20 Pages: 37-43

    • NAID

      130005056803

    • Related Report
      2014 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Animal-mounted gyroscope/accelerometer/magnetometer: in situ measurement of the movement performance of fast-start behaviour in fish2014

    • Author(s)
      Noda T, Kawabata Y, Arai N, Mitamura H, Watanabe S
    • Journal Title

      Journal of Experimental Marine Biology and Ecology

      Volume: 451 Pages: 55-68

    • Related Report
      2014 Annual Research Report
    • Peer Reviewed
  • [Presentation] Schooling behaviour of Pacific bluefin tuna in a sea net cage monitored by a novel fine-scale positioning telemetry system.2016

    • Author(s)
      Hiromichi Mitamura, Wataru Saito, Kotaro Ichikawa, Nobuaki Arai, Junichi Takagi, Ko Fujioka, Hiromu Fukuda.
    • Organizer
      21st Symposium of the International Society on Biotelemetry.
    • Place of Presentation
      Luven, Belgium
    • Year and Date
      2016-05-22
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Diurnal patterns in burrow use of the Japanese eel Anguilla japonica by using ultrasonic telemetry2016

    • Author(s)
      Kotaro Ichikawa, Hiromichi Mitamura, Nobuaki Arai, Kenta Serizawa, Shun Watanabe, Katsumi Tsukamoto.
    • Organizer
      21st Symposium of the International Society on Biotelemetry. (Luven, May 22nd-24th 2016).
    • Place of Presentation
      Luven, Belgium
    • Year and Date
      2016-05-22
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Developing an inter-individual communication biotelemetry system2016

    • Author(s)
      Minlee Yap, Yoshinori Miyamoto, Nobuaki Arai, Hiromichi Mitamura, Keiichi Uchida, Satoko Soen Kimura, Kazushi Miyashita. Hokuto Shirakawa, Yuichi Tsuda, Takashi Kitagawa.
    • Organizer
      21st Symposium of the International Society on Biotelemetry.
    • Place of Presentation
      Luven, Belgium
    • Year and Date
      2016-05-22
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得2 超音波バイオテレメトリーによる複数同時測位手法の開発.2016

    • Author(s)
      高木淳一・市川光太郎・荒井修亮・宮本佳則・内田圭一・小路 淳・三田村啓理
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得3 福井県久々子湖におけるニホンウナギの行動圏および日周行動.2016

    • Author(s)
      竹野遼馬・市川光太郎・荒井修亮・三田村啓理・高木淳一・海部健三・矢田 崇
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得4 宮崎県美郷町におけるニホンウナギの音響測位と巣穴利用特性2016

    • Author(s)
      市川光太郎・三田村啓理・荒井修亮・芹澤健太・渡邊 俊・塚本勝巳
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得5 実験地における黄ウナギの行動観察.2016

    • Author(s)
      芹澤健太・渡邊 俊・市川光太郎・三田村啓理・荒井修亮・塚本勝巳
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得6 汽水域に放流したニホンウナギ養殖個体の移動.2016

    • Author(s)
      野田琢嗣・和田敏裕・岩崎高資・佐藤太津真・成田 薫・松本育夫・堀 友彌・三田村啓理・荒井修亮
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得7  震災後の松川浦におけるマアナゴの行動・生態の解明⑤.2016

    • Author(s)
      堀 友彌・野田琢嗣・和田敏裕・岩崎高資・佐藤太津真・成田 薫・松本育夫・荒井修亮・三田村啓理
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] データ高回収率を実現するバイオロギング・システムの構築 〜平成27年度に得られたプロジェクト成果と技術的課題〜.2016

    • Author(s)
      白川北斗・宮下和士・津田裕一・北川貴士・中村乙水・宮本佳則・内田圭一・ヤップミンリー・三田村啓理・野田琢嗣・奥山隼一・木村里子・市川光太郎・荒井修亮
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] クロマグロ幼魚の群れ行動—グライド同調と体サイズ.2016

    • Author(s)
      野田琢嗣・藤岡 紘・福田漠生・三田村啓理・市川光太郎・荒井修亮
    • Organizer
      平成28年度日本水産学会春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2016-03-27
    • Related Report
      2015 Annual Research Report
  • [Presentation] Feedinghabits of Mekong giant catfish in Kaeng Krachan reservoir, Thailand.2016

    • Author(s)
      Yuichiro Tsuji, Hiromichi Mitamura, Takuji Noda, Nobuaki Arai, Hiroyuki Yamane, Yasushi Mitsunaga, Thavee Viputhanumas
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Development of the pinger with an accelerometer for classification of feeding behavior of fish2016

    • Author(s)
      Jun Horie, Yusuke Ina, Hiromichi Mitamura, Kazuyuki Moriya, Nobuaki Arai, Toyoki Sasakura
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Behavior of juvenile yellowfin tuna responding to light-dark change under an open sea net cage environment.2016

    • Author(s)
      Yusuke Komoto, Takuji Noda, Hiromu Fukuda, Ko Fujioka, Junichi Takagi, Kotaro Ichikawa, Nobuaki Arai, Hiromichi Mitamura
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Factors affecting neighbor distance of a school of 0-aged bluefin tuna2016

    • Author(s)
      Wataru Saito, Hiromichi Mitamura, Kotaro Ichikawa, Nobuaki Arai, Junichi Takagi, Ko Fujioka, Hiromu Fukuda
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Monitoring of Japanese eel Anguilla japonica by fine-scale positioning system in the Lake Kugushi, Fukui, Japan2016

    • Author(s)
      Ryoma Takeno, Kotaro Ichikawa, Nobuaki Arai, Hiromichi Mitamura, Junichi Takagi, Kenzo Kaifu, Takashi Yada
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Application of axis-free acceleration data for the classification of feeding behavior of sit-and-wait fish.2016

    • Author(s)
      Yusuke Ina, Jun Horie, Takuji Noda, Nobuaki Arai, Hiromichi Mitamura
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Evaluation of multi-path effects on depth detection provided by acoustic transmitters in shallow water2016

    • Author(s)
      Junichi Takagi, Kotaro Ichikawa, Nobuaki Arai, Yoshinori Miyamoto, Keiichi Uchida,Ko Fujioka, Hiromu Fukuda, Hiromichi Mitamura
    • Organizer
      The 4th Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2016-03-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Endangered species Mekong giant catfish: Acoustic telemetry solves the mystery of the giant catfish ecology.2015

    • Author(s)
      Hiromichi Mitamura.
    • Organizer
      The Ninth Annual Meeting of Asian Fisheries Acoustics Society.
    • Place of Presentation
      Bangkok, Thailand
    • Year and Date
      2015-11-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research / Invited
  • [Presentation] Developing an Inter-individual communication biologging system.2015

    • Author(s)
      Minlee Yap, Yoshinori Miyamoto, Keiichi Uchida, Miyashita Kazushi, Hokuto Shirakawa, Yuichi Tsuda, Nobuaki Arai, Hiromichi Mitamura, Satoko Soen Kimura.
    • Organizer
      The Ninth Annual Meeting of Asian Fisheries Acoustics Society.
    • Place of Presentation
      Bangkok, Thailand
    • Year and Date
      2015-11-16
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Spatio-temporal distribution of the white-spotted conger on tsunami-stricken fishing ground in Fukushima2015

    • Author(s)
      Tomoya Hori, Takuji Noda, Toshihiro Wada, Takashi Iwasaki, Hiromichi Mitamura, and Nobuaki Arai.
    • Organizer
      3rd Annual International Conference on Fish Telemetry
    • Place of Presentation
      Halifax, Canada
    • Year and Date
      2015-07-13
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Regular behaviour and movement patterns in snow crabs at dark continental shelf2015

    • Author(s)
      Hiromichi Mitamura, Tetsuya Ohtani, Tameo Ozaki, Nao Yoshida, Nobuaki Arai.
    • Organizer
      3rd Annual International Conference on Fish Telemetry
    • Place of Presentation
      Halifax, Canada
    • Year and Date
      2015-07-13
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] Construction of advanced biologging systems for high rates of data-recovery - a challenging study to clarify the dynamics of fish populations and communities-2015

    • Author(s)
      Hiromichi Mitamura, Yoshinori Miyamoto, Nobuaki Arai, Takashi Kitagawa, Keiichi Uchida, Hokuto Shirakawa, Yuichi Tsuda, Itsumi Nakamura, Takuji Noda, Kotaro Ichikawa, Junichi Okuyama, Kazushi Miyashita.
    • Organizer
      3rd Annual International Conference on Fish Telemetry
    • Place of Presentation
      Halifax, Canada
    • Year and Date
      2015-07-13
    • Related Report
      2015 Annual Research Report
    • Int'l Joint Research
  • [Presentation] 半自然環境下でのクロマグロ0歳魚の群れの空間的特徴およびグライド同調.2015

    • Author(s)
      高本悠介・野田琢嗣・福田漠生・藤岡紘・市川光太郎・荒井修亮・三田村啓理
    • Organizer
      海洋理工学会平成27年度春季大会
    • Place of Presentation
      東京海洋大、東京都港区
    • Year and Date
      2015-05-26
    • Related Report
      2015 Annual Research Report
  • [Presentation] Spatio-temporal distribution of the white-spotted conger on tsunami-stricken fishing ground in Fukushima2015

    • Author(s)
      Tomoya Hori, Takuji Noda, Toshihiro Wada, Takashi Iwasaki, Hiromichi Mitamura & Nobuaki Ara
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Habitat use and diel movement patterns of hatchery-reared Mekong giant catfish in Kaeng Krachan reservoir, Thailand2015

    • Author(s)
      Yuichiro Tsuji, Hiromichi Mitamura, Takuji Noda, Nao Yoshida, Nobuaki Arai, Hiroyuki Yamane, Yasushi Mitsunaga & Thavee Vipthanumas
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Fine-scale monitoring of Adélie penguins commuting pathways in a fast sea-ice area2015

    • Author(s)
      Junichi Takagi, Hiromichi Mitamura, Takuji Noda & Akinori Takahashi
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Swimming locomotion of terrestrial snakes, Elaphe quadrivirgata and Elaphe climacophora2015

    • Author(s)
      Runa Tabata, Hideaki Nishizawa, Tomoya Hori, Hiromichi Mitamura & Nobuaki Arai
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Glide synchronization and intra-school spatial movement in schooling 0-aged Pacific bluefin tunas under a semi-natural environment2015

    • Author(s)
      Yusuke Komoto, Takuji Noda, Hiromu Fukuda, Ko Fujioka, Kotaro Ichikawa, Nobuaki Arai & Hiromichi Mitamura
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] The tracking of giant freshwater stingrays (Himantura chaophraya) using acoustic telemetry in the Chao Phya River, Thailand2015

    • Author(s)
      Nao Yoshida, Hiromichi Mitamura, Takuji Noda, Nobuaki Arai, Hiroyuki Yamane, Yasushi Mitsunaga & Thavee Viputhanumas
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Identification of feeding behaviors in sit-and-wait fish from axis-free motion data2015

    • Author(s)
      Yuichiro Mashino, Yusuke Ina, Takuji Noda, Nobuaki Arai & Hiromichi Mitamura
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Development of event-driven acoustic transmitter for monitoring behavior of aquatic animals2015

    • Author(s)
      Yusuke Ina, Yuichiro Mashino, Takuji Noda, Nobuaki Arai & Hiromichi Mitamura
    • Organizer
      The 3rd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      京都大学、京都府京都市
    • Year and Date
      2015-03-15
    • Related Report
      2014 Annual Research Report
  • [Presentation] Evaluation of positioning accuracy using frequency-modulated tones for biotelemetry2014

    • Author(s)
      Masahiro Nakagawa, Kotaro Ichikawa, Toyoki Sasakura, Tomio Shinke, Hiromichi Mitamura, Kazuyuki Moriya, Nobuaki Arai
    • Organizer
      The 2nd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      Kyoto, Japan
    • Related Report
      2013 Annual Research Report
  • [Presentation] Development of a fine scale acoustic positioning and telemetry system for schooling behavior of aquatic animals2014

    • Author(s)
      Kazunori Kikuchi, Nobuaki Arai, Hiromichi Mitamura, Kotaro Ichikawa, Ko Fujioka, Hiromu Fukuda
    • Organizer
      The 2nd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      Kyoto, Japan
    • Related Report
      2013 Annual Research Report
  • [Presentation] A study on two attachment methods of tags and acoustic devices for monitoring the behavior of Japanese common sea cucumber Apostichopus japonicus2014

    • Author(s)
      Takumi Fujino, Hiromichi Mitamura, Nobuaki Arai, Reiji Masuda, Hideki Sawada, Kotaro Ichikawa
    • Organizer
      The 2nd Design Symposium on Conservation of Ecosystem (SEASTAR2000)
    • Place of Presentation
      Kyoto, Japan
    • Related Report
      2013 Annual Research Report
  • [Presentation] バイオロギングによる水圏生物の行動情報の取得 4 超音波発信機を用いたマナマコへのテレメトリー手法の開発2014

    • Author(s)
      藤野 匠、澤田英樹、益田玲爾、市川光太朗、三田村啓理、荒井修亮
    • Organizer
      平成26年度日本水産学会春季大会
    • Place of Presentation
      函館、日本
    • Related Report
      2013 Annual Research Report
  • [Presentation] 超音波バイオテレメトリーのための周波数変調方式を用いた超音波発信機の開発と評価2013

    • Author(s)
      中川正博、市川光太郎、笹倉豊喜、新家富雄、荒井修亮、三田村啓理
    • Organizer
      海洋理工学会平成25年度春季大会
    • Place of Presentation
      東京、日本
    • Related Report
      2013 Annual Research Report

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Published: 2013-05-21   Modified: 2019-07-29  

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