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Excavation of the late Miocene primate fauna in Nakali, Kenya and its evolutionary implications

Research Project

Project/Area Number 22255006
Research Category

Grant-in-Aid for Scientific Research (A)

Allocation TypeSingle-year Grants
Section海外学術
Research Field Physical anthropology
Research InstitutionKyoto University

Principal Investigator

NAKATSUKASA Masato  京都大学, 理学(系)研究科(研究院), 教授 (00227828)

Co-Investigator(Kenkyū-buntansha) KUNIMARSU Yutaka  京都大学, 大学院理学研究科, 准教授 (80243111)
OGIHARA Naomichi  慶応義塾大学, 理工学部, 准教授 (70324605)
SAKAI Tetsuya  島根大学, 総合理工学部, 准教授 (90303809)
NAKATANI Hideo  鹿児島大学, 大学院理工学研究科, 教授 (20180424)
SANEYOSHI Mototaka  株式会社林原生物化学研究所, 類人猿研究センター, 研究員 (50522140)
石田 英實  聖泉大学, 看護学部, 教授 (60027480)
Project Period (FY) 2010-04-01 – 2014-03-31
Project Status Completed (Fiscal Year 2013)
Budget Amount *help
¥37,830,000 (Direct Cost: ¥29,100,000、Indirect Cost: ¥8,730,000)
Fiscal Year 2012: ¥11,960,000 (Direct Cost: ¥9,200,000、Indirect Cost: ¥2,760,000)
Fiscal Year 2011: ¥11,960,000 (Direct Cost: ¥9,200,000、Indirect Cost: ¥2,760,000)
Fiscal Year 2010: ¥13,910,000 (Direct Cost: ¥10,700,000、Indirect Cost: ¥3,210,000)
Keywords古人類学 / 東アフリカ / ケニア / 中新世 / 化石 / 霊長類 / 古環境 / 大型類人猿 / ヒト上科 / オナガザル科 / 進化
Research Abstract

We conducted paleontological and geological survey of the Nakali Formation (9.8-9.9 million years ago), documented the mammal community of this age including several primate species, and revealed its paleoenvironment and their ecological adaptations. Mammalian taxa counted 6 orders and more than 30 species. The most abundant primate is Microcolobus sp. This species is a small colobine that is comparable to the smallest extant colobine in size and was adapted for living in forested environment. The presumed forested (C3) paleoenvironment was also supported from stable carbon isotope study of fossilized enamel of herbivorous mammals. This study also revealed that C4 environmental expansion progressed after the age of the Nakali Formation in East Africa. The geological survey revealed that the Nakali Formation expands southward more widely that our previous expectation.

Report

(4 results)
  • 2013 Final Research Report ( PDF )
  • 2012 Annual Research Report
  • 2011 Annual Research Report
  • 2010 Annual Research Report
  • Research Products

    (57 results)

All 2013 2012 2011 2010 Other

All Journal Article (14 results) (of which Peer Reviewed: 13 results) Presentation (40 results) (of which Invited: 2 results) Remarks (3 results)

  • [Journal Article] Planar covariation of limb elevation angles during bipedal walking in the Japanese macaque2012

    • Author(s)
      N. Ogihara, T. Kikuchi, Y. Ishiguro, H. Makishima, M. Nakatsukasa
    • Journal Title

      I R. Soc. Interface

      Volume: Vol.9, No.74 Pages: 2180-2190

    • URL

      http://rsif.royalsocietypublishing.org/content/early/2012/03/19/rsif.2012.0026.abstract

    • Related Report
      2013 Final Research Report
    • Peer Reviewed
  • [Journal Article] アフリカの中新世旧世界ザルの進化:現生ヒト上科進化への影響2012

    • Author(s)
      中務真人他
    • Journal Title

      Anthropological Science(Japanese Series)

      Volume: Vol.120, No.2 Pages: 99-119

    • NAID

      10031145908

    • URL

      https://www.jstage.jst.go.jp/article/asj/120/2/120_121022/_article/-char/ja

    • Related Report
      2013 Final Research Report
    • Peer Reviewed
  • [Journal Article] Hind limb of the Nacholapithecus kerioi holotype and implications for its positional behavior2012

    • Author(s)
      M. Nakatsukasa, Y. Kunimatsu, et al
    • Journal Title

      Anthropological Science

      Volume: Vol. 120, No.3 Pages: 235-250

    • NAID

      10031145907

    • URL

      https://www.jstage.jst.go.jp/article/ase/advpub/0/advpub_120731/_article

    • Related Report
      2013 Final Research Report
    • Peer Reviewed
  • [Journal Article] Functional morphology and anatomy of cervical vertebrae in Nacholapithecus kerioi, a middle Miocene hominoid from Kenya2012

    • Author(s)
      Y. Kikuchi, Y. Nakano, M. Nakatsukasa, et al
    • Journal Title

      Journal of Human Evolution

      Volume: Vol.62, No.6 Pages: 677-695

    • URL

      http://www.sciencedirect.com/science/article/pii/S0047248412000413

    • Related Report
      2013 Final Research Report
    • Peer Reviewed
  • [Journal Article] Evolution of the Miocene Old World monkeys in Africa: influence on the evolution of the extant hominoids2012

    • Author(s)
      中務真人
    • Journal Title

      Anthropological Science (Japanese Series)

      Volume: 120 Issue: 2 Pages: 99-119

    • DOI

      10.1537/asj.121022

    • NAID

      10031145908

    • ISSN
      1344-3992, 1348-8813
    • Related Report
      2012 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Hind limb of the <i>Nacholapithecus kerioi</i> holotype and implications for its positional behavior2012

    • Author(s)
      Nakatsukasa M Kunimatsu Y Shimizu D Nakano Y Kikuchi Y Ishida H
    • Journal Title

      Anthropological Science

      Volume: 120 Issue: 3 Pages: 235-250

    • DOI

      10.1537/ase.120731

    • NAID

      10031145907

    • ISSN
      0918-7960, 1348-8570
    • Related Report
      2012 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Functional morphology and anatomy of cervical vertebrae in Nacholapithecus kerioi, a middle Miocene hominoid from Kenya2012

    • Author(s)
      Kikuchi Y Nakano Y Nakatsukasa M Kunimatsu Y Shimizu D Ogihara N Tsujikawa H Takano T Ishida H
    • Journal Title

      Journal of Human Evolution

      Volume: 62 Issue: 6 Pages: 677-695

    • DOI

      10.1016/j.jhevol.2012.03.002

    • Related Report
      2012 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Planar covariation of limb elevation angles during bipedal walking in the Japanese macaque2012

    • Author(s)
      N.Ogihara, T.Kikuchi, Y.Ishiguro, H.Makishima, M.Nakatsukasa
    • Journal Title

      J.R.Soc.Interface

      Volume: (印刷中) Issue: 74 Pages: 2181-2190

    • DOI

      10.1098/rsif.2012.0026

    • Related Report
      2011 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Late Miocene to Pliocene carbon isotope record of differential diet change among East African herbivores2011

    • Author(s)
      K. Uno, T. Cerling, J. Harris, Y. Kunimatsu, M. Leakey, M. Nakatsukasa, H. Nakaya
    • Journal Title

      Proceedings of the National Academy of Science of the United States of America,potsA

      Volume: Vol.108, No.16 Pages: 6509-6514

    • URL

      http://www.pnas.org/content/108/16/6509.short

    • Related Report
      2013 Final Research Report
  • [Journal Article] Forward dynamic simulation of bipedal walking in the Japanese macaque: investigation of causal relationships among limb kinematics, speed, and energetics of bipedal locomotion in a non-human primate2011

    • Author(s)
      N. Ogihara, H. Makishima, M. Nakatsukasa
    • Journal Title

      American Journal of Physical Anthropology

      Volume: Vol. 145, No. 4 Pages: 568-580

    • URL

      http://www.ncbi.nlm.nih.gov/pubmed/21590751

    • Related Report
      2013 Final Research Report
    • Peer Reviewed
  • [Journal Article] Late Miocene to Pliocene carbon isotope record of differential diet change among East African herbivores2011

    • Author(s)
      K.Uno, T.Cerling, J.Harris, Y.Kunimatsu, M.Leakey, M.Nakatsukasa, H.Nakaya
    • Journal Title

      Proc.Natl.Acad.Sci.U.S.A.

      Volume: 108 Issue: 16 Pages: 6509-6514

    • DOI

      10.1073/pnas.1018435108

    • Related Report
      2011 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Forward dynamic simulation of bipedal walking in the Japanese macaque : investigation of causal relationships among limb kinematics, speed, and energetics of bipedal locomotion in a nonhuman primate2011

    • Author(s)
      N.Ogihara, S.Aoi, Y.Sugimoto, K.Tsuchiya, M.Nakatsukasa
    • Journal Title

      Am.J.Phys.Anthropol

      Volume: 145 Issue: 4 Pages: 568-580

    • DOI

      10.1002/ajpa.21537

    • Related Report
      2011 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Late Miocene to Pliocene carbon isotope record of differential diet change among East African herbivores2011

    • Author(s)
      K.Uno, T.Cerling, J.Harris, Y.Kunimatsu, M.Leakey, M.Nakatsukasa, H.Nakaya
    • Journal Title

      Proceedings of the National Academy of Science of the United States of America

      Volume: (in press)

    • Related Report
      2010 Annual Research Report
    • Peer Reviewed
  • [Journal Article] Forward dynamic simulation of bipedal walking in the Japanese macaque : investigation of causal relationships among limb kinematics, speed, and energetics of bipedal locomotion in a non-human primate2011

    • Author(s)
      N.Ogihara, H.Makishima, M.Nakatsukasa
    • Journal Title

      American Journal of Physical Anthropology

      Volume: (in press)

    • Related Report
      2010 Annual Research Report
    • Peer Reviewed
  • [Presentation] ケニア北部の上部中新統ナカリ層およびナムルングレ層産Chilotheridium(奇蹄目,サイ科)化石2013

    • Author(s)
      半田直人,仲谷英夫 他
    • Organizer
      古生物学会第162回例会
    • Place of Presentation
      横浜国立大学海洋研究開発機構(神奈川県)
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニア北部の上部中新統ナカリ層およびナムルングレ層産Chilotheridium (奇蹄目,サイ科) 化石2013

    • Author(s)
      半田直人
    • Organizer
      古生物学会第162回例会
    • Place of Presentation
      横浜国立大学
    • Related Report
      2012 Annual Research Report
  • [Presentation] ケニア北部産、中期中新世の類人猿Nacholapithecus kerioiにおける腰椎形態2013

    • Author(s)
      菊池泰弘
    • Organizer
      第118回日本解剖学会総会・全国学術集会
    • Place of Presentation
      サンポートホール高松かがわ国際会議場
    • Related Report
      2012 Annual Research Report
  • [Presentation] 中新世類人猿の進化2012

    • Author(s)
      中務真人
    • Organizer
      第59回プリマーテス研究会
    • Place of Presentation
      日本モンキーセンター(愛知県
    • Year and Date
      2012-12-08
    • Related Report
      2013 Final Research Report
    • Invited
  • [Presentation] 後期中新世ケニア、ナカリにおける最近の調査成果について2012

    • Author(s)
      中務真人 他
    • Organizer
      第66回日本人類学会大会
    • Place of Presentation
      慶慮義塾大学(神奈川県)
    • Year and Date
      2012-11-04
    • Related Report
      2013 Final Research Report
  • [Presentation] ナカリ出土の類人猿足根骨について2012

    • Author(s)
      中務真人他
    • Organizer
      第28回日本霊長類学会大会
    • Place of Presentation
      椙山女学園大学(愛知県)
    • Year and Date
      2012-07-09
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニヤ北部ナチョラから見つかった中期中新世の原猿化石について2012

    • Author(s)
      國松豊 他
    • Organizer
      第28回日本霊長類学会大会
    • Place of Presentation
      椙山女学園大学(愛知県)
    • Year and Date
      2012-07-09
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニア・リフトバレーの後期中新世哺乳類動物相とその古環境2012

    • Author(s)
      仲谷英夫他
    • Organizer
      日本アフリカ学会第48回学術大会
    • Place of Presentation
      弘前大学(青森県)
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニア北部のナムルングレ層およびナカリ層産後期中新世サイ科(奇蹄目,哺乳綱)化石2012

    • Author(s)
      半田直人,仲谷英夫他
    • Organizer
      日本アフリカ学会第48回学術大会
    • Place of Presentation
      弘前大学(青森県)
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニア・リフトバレーのサンブルヒルズとナカリの後期中新世哺乳類動物相とその古環境2012

    • Author(s)
      仲谷英夫他
    • Organizer
      日本地球惑星科学連合2012年連合大会
    • Place of Presentation
      幕張メッセ国際会議場(千葉県)
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニア北部のナムルングレ層およびナカリ層産後期中新世エラスモテリウム族(サイ科,奇蹄目)化石2012

    • Author(s)
      半田直人,仲谷英夫他
    • Organizer
      日本地球惑星科学連合2012年連合大会
    • Place of Presentation
      幕張メッセ国際会議場(千葉県)
    • Related Report
      2013 Final Research Report
  • [Presentation] ケニア・リフトバレーの後期中新世哺乳類動物相とその古環境2012

    • Author(s)
      仲谷英夫
    • Organizer
      日本アフリカ学会第48回学術大会
    • Place of Presentation
      弘前大学
    • Related Report
      2012 Annual Research Report
  • [Presentation] ケニア北部のナムルングレ層およびナカリ層産後期中新世サイ科 (奇蹄目,哺乳綱) 化石2012

    • Author(s)
      半田直人
    • Organizer
      日本アフリカ学会第48回学術大会
    • Place of Presentation
      弘前大学
    • Related Report
      2012 Annual Research Report
  • [Presentation] ケニア・リフトバレーのサンブルヒルズとナカリの後期中新世哺乳類動物相とその古環境2012

    • Author(s)
      仲谷英夫
    • Organizer
      日本地球惑星科学連合2012年連合大会
    • Place of Presentation
      幕張メッセ国際会議場
    • Related Report
      2012 Annual Research Report
  • [Presentation] ケニア北部のナムルングレ層およびナカリ層産後期中新世エラスモテリウム族(サイ科,奇蹄目) 化石2012

    • Author(s)
      半田直人
    • Organizer
      日本地球惑星科学連合2012年連合大会
    • Place of Presentation
      幕張メッセ国際会議場
    • Related Report
      2012 Annual Research Report
  • [Presentation] ナカリ出土の類人猿足根骨について2012

    • Author(s)
      中務真人
    • Organizer
      第28回日本霊長類学会大会
    • Place of Presentation
      椙山女学園大学(愛知県)
    • Related Report
      2012 Annual Research Report
  • [Presentation] ケニヤ北部ナチョラから見つかった中期中新世の原猿化石について2012

    • Author(s)
      國松豊
    • Organizer
      第28回日本霊長類学会大会
    • Place of Presentation
      椙山女学園大学(愛知県)
    • Related Report
      2012 Annual Research Report
  • [Presentation] 後期中新世ケニア、ナカリにおける最近の調査成果について2012

    • Author(s)
      中務真人
    • Organizer
      第66回日本人類学会大会
    • Place of Presentation
      慶應義塾大学
    • Related Report
      2012 Annual Research Report
  • [Presentation] ケニア北部産・中期中新世の類人猿 Nacholapithecus kerioi における下部胸椎形態2012

    • Author(s)
      菊池泰弘
    • Organizer
      第66回日本人類学会大会
    • Place of Presentation
      慶應義塾大学
    • Related Report
      2012 Annual Research Report
  • [Presentation] 中新世類人猿の進化2012

    • Author(s)
      中務真人
    • Organizer
      第59回プリマーテス研究会
    • Place of Presentation
      日本モンキーセンター
    • Related Report
      2012 Annual Research Report
    • Invited
  • [Presentation] Recent advances in the study of African Miocene apes and monkeys2011

    • Author(s)
      Y. Kunimatsu, M. Nakatsukasa, 他
    • Organizer
      World Conference on Paleontology and Stratigraphy (WCPS2011)
    • Place of Presentation
      Nakhon Ratchasima (Thailand)
    • Year and Date
      2011-11-29
    • Related Report
      2013 Final Research Report 2011 Annual Research Report
  • [Presentation] Paleoenvironment of Nakali inferred from extant colobine ecology2011

    • Author(s)
      中務真人,國松 豊,仲谷英夫,実吉玄置,酒井哲弥
    • Organizer
      第65回日本人類学会大会
    • Place of Presentation
      沖縄県立博物館・美術館(沖縄県)
    • Year and Date
      2011-11-29
    • Related Report
      2013 Final Research Report 2011 Annual Research Report
  • [Presentation] ケニヤ共和国ナカリ地域出土の多様な後期中新世の狭鼻猿相2011

    • Author(s)
      國松 豊,中務真人他
    • Organizer
      第65回日本人類学会大会
    • Place of Presentation
      沖縄県立博物館・美術館(沖縄県)
    • Year and Date
      2011-11-29
    • Related Report
      2013 Final Research Report 2011 Annual Research Report
  • [Presentation] Diverse catarrhine fauna of the early Late Miocene of East Africa: evidence from Nakali, north-central Kenya2011

    • Author(s)
      Y. Kunimatsu, M. Nakatsukasa, 他
    • Organizer
      East African Association for Paleoanthropology and Paleontology: 3rd Conference
    • Place of Presentation
      Addis Ababa (Ethiopia)
    • Year and Date
      2011-08-09
    • Related Report
      2013 Final Research Report
  • [Presentation] Diverse catarrhine fauna of the early Late Miocene of East Africa : evidence from Nakali, north-central Kenya2011

    • Author(s)
      Y.Kunimatsu, M.Nakatsukasa, D.Shimizu, A.Yamamoto, T.Sakai, H.Nakaya, 他
    • Organizer
      East African Association for Paleoanthropology and Paleontology : 3rd Conference
    • Place of Presentation
      Addis Ababa (Ethiopia)
    • Year and Date
      2011-08-09
    • Related Report
      2011 Annual Research Report
  • [Presentation] ナカリにおける発掘調査と初期コロブス類進化についての新知見2011

    • Author(s)
      中務真人,國松 豊,仲谷英夫,実吉玄旦,酒井哲弥
    • Organizer
      第27回日本霊長類学術大会
    • Place of Presentation
      犬山国際観光センターフロイデ(愛知県)
    • Year and Date
      2011-07-16
    • Related Report
      2013 Final Research Report 2011 Annual Research Report
  • [Presentation] アフリカ中期中新世の化石類人猿Nacholapithecusにおける弓下窩の消失2011

    • Author(s)
      國松豊, 中務真人, 清水大輔, 辻川寛, 中野良彦, 荻原直道, 菊池泰弘, 石田英實
    • Organizer
      第27回日本霊長類学術大会
    • Place of Presentation
      犬山国際観光センターフロイデ(愛知県)
    • Year and Date
      2011-07-16
    • Related Report
      2011 Annual Research Report
  • [Presentation] 初期ヒト科の体重性差を巡る議論について2010

    • Author(s)
      中務真人
    • Organizer
      第64回日本人類学会大会
    • Place of Presentation
      だて歴史の杜カルチャーセンター(北海道)
    • Year and Date
      2010-10-02
    • Related Report
      2013 Final Research Report 2010 Annual Research Report
  • [Presentation] 仙尾部骨格形態によるマカクの尾長推定2010

    • Author(s)
      東島沙弥佳, 中務真人
    • Organizer
      第64回日本人類学会大会
    • Place of Presentation
      だて歴史の杜カルチャーセンター(北海道)
    • Year and Date
      2010-10-02
    • Related Report
      2010 Annual Research Report
  • [Presentation] ヒトと類人猿の胸郭形態2010

    • Author(s)
      加賀谷美幸, 荻原直道, 中務真人
    • Organizer
      第64回日本人類学会大会
    • Place of Presentation
      だて歴史の杜カルチャーセンター(北海道)
    • Year and Date
      2010-10-02
    • Related Report
      2010 Annual Research Report
  • [Presentation] 炭素同位体分析による稲荷山貝塚縄文人の食性と抜歯形式の関係の解明2010

    • Author(s)
      日下宗一郎, Kevin T, UNO, Thure E.CERLING, 中務真人
    • Organizer
      第64回日本人類学会大会
    • Place of Presentation
      だて歴史の杜カルチャーセンター(北海道)
    • Year and Date
      2010-10-01
    • Related Report
      2010 Annual Research Report
  • [Presentation] Archaic postcranium of middle Miocene apes and parallel evolution of modern postcranial anatomy in extant apes2010

    • Author(s)
      M. Nakatsukasa
    • Organizer
      International Primatological Society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-17
    • Related Report
      2013 Final Research Report 2010 Annual Research Report
  • [Presentation] Fossil catarrhines from the early late Miocene of Nakali in Kenya2010

    • Author(s)
      Y. Kunimatsu, M. Nakatsukasa, 他
    • Organizer
      International Primatological Society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-13
    • Related Report
      2013 Final Research Report 2010 Annual Research Report
  • [Presentation] Diet analysis for late Miocene catarrhines from Nakali in Kenya2010

    • Author(s)
      D. Shimizu, T. Ikarashi, Y. Kunimatsu, M. Nakatsukasa, E. Mbua
    • Organizer
      International Primatological Society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-13
    • Related Report
      2013 Final Research Report
  • [Presentation] Late Miocene Paleoenvironments of Hominoidsmesowear analysis of fossil ungulate cheek teeth from northern Kenya2010

    • Author(s)
      H. Nakaya, K. Uno, A. Fukuchi, Y. Kunimatsu, M. Nakatsukasa, 他
    • Organizer
      International Primatological Society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-13
    • Related Report
      2013 Final Research Report
  • [Presentation] Uniqueness of thoracic cage morphology in extant hominoids : implications for the evolution of positional behaviors.2010

    • Author(s)
      M.Kagaya, N.Ogihara, M.Nakatsukasa, B.Senut
    • Organizer
      International Primatological society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-13
    • Related Report
      2010 Annual Research Report
  • [Presentation] Diet analysis for late Miocene catarrhines from Nakali in Kenya.2010

    • Author(s)
      D.Shimizu, T.Ikarashi, Y.Kunimatsu, M.Nakatsukasa, E.Mbua
    • Organizer
      International Primatological Society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-13
    • Related Report
      2010 Annual Research Report
  • [Presentation] Late Miocene Paleoenvironments of Hominoids-mesowear analysis of fossil ungulate cheek teeth from northern Kenya.2010

    • Author(s)
      H.Nakaya, K.Uno, A.Fukuchi, Y.Kunimatsu, M.Nakatsukasa, 他4名
    • Organizer
      International Primatological society XXIII Congress
    • Place of Presentation
      京都大学(京都府)
    • Year and Date
      2010-09-13
    • Related Report
      2010 Annual Research Report
  • [Presentation] Evolution of locomotor adaptation in primates2010

    • Author(s)
      M. Nakatsukasa
    • Organizer
      International Symposium on Application of Biomechanical Control Systems to Precision Engineering
    • Place of Presentation
      アクアマリンふくしま(福島県)
    • Year and Date
      2010-07-22
    • Related Report
      2013 Final Research Report
  • [Presentation] Evolution of locomotor adaptation in primates2010

    • Author(s)
      M.Nakatsukasa
    • Organizer
      International Symposium on Application of Biomechanical Control Systems to Precision Engineering
    • Place of Presentation
      アクアマリンふくしま(福島県)(招待講演)
    • Year and Date
      2010-07-22
    • Related Report
      2010 Annual Research Report
  • [Remarks]

    • URL

      http://anthro.zool.kyoto-u.ac.jp/research/index.html

    • Related Report
      2013 Final Research Report
  • [Remarks]

    • URL

      http://anthro.zool.kyoto-u.ac.jp/report/index.html

    • Related Report
      2011 Annual Research Report
  • [Remarks]

    • URL

      http://anthro.zool.kyoto-u.ac.jp/research/index.html

    • Related Report
      2010 Annual Research Report

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Published: 2010-08-23   Modified: 2019-07-29  

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